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D 1.2 Report on the impact of current and historical habitat fragmentation on biodiversity

D 2.2 Report on meta-analysis of selected Natura 2000 sites [PDF, 838 KB]

D 3.2 Report on historic effect of land use changes based on organism and habitat [PDF, 656 KB]

D 3.3 Report on GIS database and identification of threatened habitats [PDF, 382 KB]

D 4.1-4.3 Exploring European land use change scenarios: thematic downscaling and regional variability [PDF, 8.87 MB]

D 4.4 Report on results of land use projections [PDF, 4.53 MB]

D 5.2 Report in policy options of biodiversity management [PDF, 787 KB]

D 6.3-6.4 Lists of published outputs of project; scientific and non-scientific paper contributions (see below).


1.1 Published in international scientific journals

Dover, J. & Settele J. (2009). The influences of landscape structure on butterfly distribution and movement: a review. Journal of Insect Conservation. 13:327.

Hambäck P.A., Summerville K.S., SteffanDewenter I., Krauss J., Englund G., and Crist T.O. (2007). Habitat specialization, body size, and family identity explain lepidopteran densityarea relationships in a crosscontinental comparison. PNAS 104: 8368-8373.

Krauss, J., Alfert, T., SteffanDewenter, I. (2009). Habitat area but not habitat age determines wild bee richness in limestone quarries. Journal of Applied Ecology 46: 194-202.

Kuussaari, M., Bommarco, R., Heikkinen, R. K., Helm, A., Krauss, J., Lindborg, R., Öckinger, E., Pärtel, M., Pino, J., Rodà, F., Stefanescu, C., Teder, T., Zobel, M., SteffanDewenter, I. (2009). Extinction debt: a challenge for biodiversity conservation. Trends in Ecology & Evolution 24: 564-571.

Öckinger, E., Dannestam, Å. & Smith, H.G. (2009) The importance of fragmentation and habitat quality of urban grasslands for butterfly diversity. Landscape and Urban Planning 93(1): 31-37

Pöyry, J., Paukkunen, J., Heliölä, J. & Kuussaari, M. (2009) Relative contributions of local and regional factors to species richness and total density of butterflies and moths in seminatural grasslands. Oecologia 160:577-587.

Raatikainen, K.M., Heikkinen, R.K., Luoto M. (2009). Relative importance of habitat area, connectivity, management and local factors for vascular plants: spring ephemerals in boreal seminatural grasslands. Biodiversity Conservation 18:1067-1085.

Bommarco R., J.C. Biesmeijer, B. Meyer, S.G. Potts, J. Poyry, S.P.M. Roberts, I. Steffan-Dewenter, E. Ockinger. Dispersal capacity and diet breadth modify the response of wild bees to habitat loss. Proc. R. Soc. Lond. B 277: 2075-2082.

Krauss J., R. Bommarco, M. Guardiola, R.K. Heikkinen, A. Helm, M. Kuussaari, R. Lindborg, E. Ockinger, M. Partel, J. Pino, J. Poyry, K.M. Raatikainen, A. Sang, C. Stefanescu, T. Teder, M. Zobel & I. Steffan-Dewenter (2010). Habitat fragmentation causes immediate and time-delayed biodiversity loss at different trophic levels. Ecology Letters 13: 597-605.

Schweiger O., Biesmeijer J.C., Bommarco R., Hickler T., Hulme P.E., Klotz S., Kuhn I., Moora M., Nielsen A., Ohlemuller R., Petanidou Th., Potts S.G., Pysek P., Stout J.C., Sykes M.T., Tscheulin Th., Vila M., Walther G-R., Westphal C., Winter M., Zobel M. & Settele J. 2010. Multiple stressors on biotic interactions: how climate change and alien species interact to affect pollination. Biological reviews.

Johan Ekroos, Janne Heliola & Mikko Kuussaari (2010) Homogenization of lepidopteran communities in intensively cultivated agricultural landscapes. Journal of Applied Ecology 47: 459-467.

Ockinger, E., Schweiger, O., Crist, T.O., Debinski, D.M., Krauss, J., Kuussaari, M., Petersen, J.D., Poyry, J., Settele, J., Summerville, K.S. & Bommarco, R. (2010): Life-history traits predict species responses to habitat area and isolation – A cross-continental synthesis. Ecology Letters 13: 969-979.

Hamback, P.A., Bergman, K.-O., Bommarco, R., Krauss, J., Kuussaari, M., Poyry, J., Ockinger, E. (2010) Allometric density responses in butterflies: the response to small and large patches by small and large species. Ecography.

Helm A., Oja T., Saar L., Takkis K., Talve T. & Partel M. 2009. Human influence lowers plant genetic diversity in communities with extinction debt. Journal of Ecology 97(6):1329-1336.

Bruckmann S.V., Krauss J. & Steffan-Dewenter I. 2010. Butterfly and plant specialists suffer from reduced connectivity in fragmented landscapes. Journal of Applied Ecology, 47: 799–809.

Ockinger E., K-O. Bergman, M. Franzen, T. Kadlec, J. Krauss, M. Kuussaari, J. Poyry, H.G. Smith, I. Steffan-Dewenter, R. Bommarco. (2012) The landscape matrix modifies the effect of habitat fragmentation in grassland butterflies. Landscape ecology 27: 121–131.

Lindborg, R., Helm, A., Bommarco, R., Heikkinen, R.K., Kuhn, I., Pykala, J. & Partel, M. (2011) Effect of habitat area and isolation on plant trait distribution in European forests and grasslands. Ecography in press.

Ockinger E., R. Lindborg, N.E. Sjodin & R. Bommarco. (2011) Landscape matrix modifies species richness of plants and insects in grassland fragments. Ecography 34: 001–009.

Marini, L., Bruun, H. H., Heikkinen, R. K., Helm, A., Honnay, O., Krauss, J., Kuhn, I., Lindborg, R., Partel, M. & Bommarco, R. (2012) Traits related to species persistence and dispersal explain changes in plant communities subjected to habitat loss. Diversity and Distributions, DOI: 10.1111/j.1472-4642.2012.00893.x.

1.2 Other national and international publications

Bommarco R., E. Öckinger & A. Helm. (2009) Life History Traits in Insects and Habitat Fragmentation. Chapter in Atlas of Biodiversity Risk.

Brückmann, S., Krauss, J., SteffanDewenter, I. (2008): Habitat connectivity affects butterfly diversity and community composition in calcareous grasslands. Verhandlungen der Gesellschaft für Ökologie 38:234.

Brückmann, S., Krauss, J., SteffanDewenter, I. (2008): Impact of local and surrounding landscape predictors on butterfly diversity in seminatural grasslands. Future of Butterflies in Europe II, Wageningen, Netherlands. Page 43.

Cousins S. & Lindborg I. (2009) Biodiversity in historical landscapes (in Swedish) Miljöforskning No 2.

Krauss, J. & SteffanDewenter, I. (2008): Is time delayed extinction a major threat for grassland butterflies and vascular plants? Verhandlungen der Gesellschaft für Ökologie 38:235.

Krauss, J., Bommarco, R., SteffanDewenter, I. (2008): Extinction debt of vascular plants and butterflies in five European countries. XXIII International Congress of Entomology, Durban South Africa. Abstract No: 1025.

Vogiatzakis I.N., S.R.M. Roberts, T. Stirpe & S.G. Potts. (2009) Evaluating Changes in and Around Natura 2000 Sites: a Proposed Methodology. Chapter in Atlas of Biodiversity Risk.

Settele, J., Penev, L., Georgiev, T., Grabaum, R., Grobelnik, V., Hammen, V., Klotz, S., Kotarac, M. & I. Kuhn (Eds) (2010) Atlas of Biodiversity Risk.

The Atlas of Biodiversity Risk, a major multidisciplinary project, was published on 22nd of May. The book is first of its kind and summarizes in a comprehensive, easy-to-read and richly illustrated form the major threats of biodiversity at a global scale. The global climate and land use changes, environmental pollution loss of pollinators and biological invasions are among the major risks of biodiversity identified by this comprehensive study. The impacts and consequences of biodiversity loss are analyzed with an emphasis on the socio-economic drivers and their effect on society. Three scenarios of potential future are the baseline for predicting impacts and explore options for mitigating adverse effects at several spatio-temporal scales. The Atlas is divided into chapters which mostly deal with particular pressures. Members of the COCONUT project have wtitten a special chapter on land use changes, containing the following contributions:

  • Bommarco, R. Land Use, Its Change and Effects on Biodiversity.
  • Reginster, I., Rounsevell, M., Butler, A. & N. Dendoncker. Land Use Change Scenarios for Europe.
  • Vogiatzakis, I.N., Roberts, S.P.M., Stirpe, M.T., & S.G. Potts. Evaluating Land Use Changes in and around Natura 2000 Sites: a Proposed Methodology.
  • Bommarco, R., Ockinger, E. & A. Helm. Life History Traits in Insects and Habitat Fragmentation.
  • Nagy, L., Dendoncker, N., Butler, A., Reginster, I., Rounsevell, M., Grabherr, G., Gottfried, M., & H. Pauli. Where Have All the Flowers Gone? From Natural Vegetation to Land Use – Land Cover Types: Past Changes and Future Forecasts.
  • Bomarco, R., Future Land Use Related Challenges for Biodiversity Research and Conservation.
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